An organization may have to share the same SMTP address space between two or more different e-mail systems. For example, you may have to share the SMTP address space between Exchange and a third-party e-mail system, or between Exchange environments that are configured in different Active Directory forests. In these scenarios, users in each e-mail system have the same domain suffix as part of their e-mail addresses.
http://technet.microsoft.com/en-us/library/bb676395.aspx
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